As in mammals, a bird’s skin has sensory nerve endings that detect heat, cold, pressure, and pain. Birds don’t rely on touch as extensively as humans, but it is still an important sense to them, especially in flight. They are very sensitive to changes in air temperature, pressure, and wind speed. Those changes are transferred down the feathers to nerve endings in the skin. Filo-plumes are numerous around the bases of major flight feathers that have sensory corpuscles around the base of each feather shaft. As the remiges are moved during flight, the filo-plumes assist the bird in judging what position its flight feathers are in, and give it feedback as it shifts the position of its feathers.